What's Different About These Common Birds? Part 3

Dark-eyed Acorn Woodpeckers in Baja California Sur (B.C.S.)

The distinctive “clown face” of the Acorn Woodpecker (Melanerpes formicivorus) is familiar to most birders along the Pacific Coast of North America. This species has one of the most elongated latitudinal ranges of any woodpecker in the Americas, extending north to south-central Washington state and south to the Andes Mountains of Colombia. It is the only woodpecker species native to both North and South America. Acorn Woodpeckers are essentially non-migratory and somewhat restricted to oak-dominated woodlands. Their extensive range is broken up by expanses of desert and other plant communities devoid of oak woodlands. The non-contiguous nature of their range genetically isolates many of these populations. The Vizcaino Desert, which bisects the middle third of the Baja Peninsula, is a major barrier to non-migratory arboreal species that reside in the Cape District of B.C.S. As discussed in the first two parts of this series, a number of taxa found in B.C.S. occur nowhere else. In the case of the Acorn Woodpecker, there are many subtle and not-so-subtle geographic variations in plumage (described and illustrated below). However, among all the disjunct populations of Acorn Woodpeckers, only the birds of Baja California Sur lack the conspicuous white eye.

This male, photographed near Grants Pass, Oregon on 13 January 2007, is typical of the Acorn Woodpeckers (subspecies bairdi) found in California, Oregon, and Washington, and matches depictions of this species found in the popular North American field guides. The underside of this bird exhibits the following features: a broad and nearly solid black breast band that frames the throat and connects to a broad black “mask” surrounding the eye, and a mostly clean white lower breast, belly, undertail, and flanks. In the U.S., Acorn Woodpeckers show dense black streaking immediately below the black breast band, but virtually no streaking on the white belly and sparse narrow black streaks on the otherwise white flanks. (Photograph taken by Phil Hicks using an Olympus C750UZ camera)

This female Acorn Woodpecker, part of a small isolated population (subspecies angustifrons) in Baja California Sur (Koenig et al. 1995), was photographed near San Antonio de la Sierra, B.C.S., Mexico on 12 January 2009. We encountered 75+ Acorn Woodpeckers in this area, and all those seen well were similar in appearance to this bird. They had dark eyes and were heavily streaked along their flanks from their lower breast to their undertail coverts. Only the very middle of the belly lacked streaking. Their black flank streaks were much broader than those of birds in the U.S. I’ve seen hundreds of Acorn Woodpeckers in Oregon and California and none have shown the amount of heavy black flank streaking exhibited by this individual. The portion of the breast band that is solid black is not as broad in the B.C.S. birds and, as evidenced by this bird, they show lots of white-tipped feathers in the black breast band. Northern birds show far fewer white-tipped feathers in the breast band, giving their breast bands the appearance of being broader and more solid black. The primary difference between males and females is the head pattern. In males, the red crown patch extends farther forward and the black does not extend across the forehead, which it does in females. (Photo by David Irons using a Canon EOS XSI 450D camera and an EF 100-400mm f/4.5-5.6 lens)

Variations in plumage and eye color between the B.C.S. population (on left) and birds in the U.S. (on right) are evident in this side-by-side comparison. These close-ups were cropped from the two photos above. The most obvious difference between these birds is eye color. The dark eye (red iris) seen on the bird on the left is unique to the B.C.S. population. All other populations of Acorn Woodpeckers in the U.S., mainland Mexico, Central America, and northern South America show white irises as adults. Note also that the somewhat circular, light-colored area wrapping around the face, referred to as the “facial circle” in this discussion, is entirely yellow on the bird on the left and mostly white with just a bit of yellow on the throat of the bird on the right. Again, only the B.C.S. birds show an all yellow facial circle. There are also subtle differences in the width of the facial circle. It is a little wider between the eye and the bill and a little narrower across the forehead on the B.C.S. birds. In all other Acorn Woodpecker populations, the light-colored facial circle is predominantly white with varying amounts of yellow on the throat.

This female Acorn Woodpecker, likely the subspecies striatipectus (Koenig et al. 1995), was photographed at Chinchona, Costa Rica in February 2007. Its features are somewhat intermediate between the B.C.S. birds and the typical birds found in North America. Like all Acorns (other than those in B.C.S.), it has white irises. The underparts are more streaked than birds in the U.S., but less heavily streaked on the flanks than those of the B.C.S. population. The black breast band is similar to the B.C.S. birds, in that it is not as broad and has a lot of white spotting. However, the facial circle of this bird is mostly white, with yellow restricted to the throat. This bird shows a broader white band across the forehead compared to the rather narrow band of yellow across the forehead of the B.C.S. birds. Note that this individual shows a few red-tipped feathers at the base of the throat, where the yellow on the throat meets the top of the black breast band. This smattering of red-tipped feathers is always shown by Acorn Woodpeckers from Oaxaca, Mexico to Panama (Koenig et al. 1995), but appears on only about 5% of the U.S. specimens (subspecies bairdi and aculeata) in the San Diego Natural History Museum (SDNHM). All four of that museum’s angustifrons specimens have red-tipped breast feathers (Phil Unitt pers. comm.). I took several images of the San Antonio de la Sierra bird shown in the images above, and found only one angle where the bird appears to show a single red-tipped breast feather. (Photograph was by Bill Tice using an Olympus D-550 camera through a Zeiss Diascope 65T)

Paying attention to geographical differences in common species is a great way to maintain interest in species of birds that are otherwise ignored during a typical day of birding. While I was able to pick up several life birds in Baja, the time spent studying the birds discussed in this three-part series was, perhaps, the most interesting aspect of the trip. My post-trip research and quest to find comparison photos answered a few questions, but most importantly raised several new ones. The Birds of North America Online is a fantastic web-based resource co-sponsored by the Cornell Lab of Ornithology and the American Ornithologists’ Union (AOU). When it came to answering specific questions about subspecies, range, and plumage variations in Acorn Woodpeckers, I referenced it often. I sincerely thank Phil Hicks, Sylvia Maulding, Ryan Merrill, and Bill Tice for allowing me to use their excellent photos, and I am also appreciative of those who provided photos that I examined, but did not include in this photo essay. I am indebted and grateful to Phil Unitt, curator of the San Diego Natural History Museum, who took time from his busy schedule to examine study skins and offer details about the various Acorn Woodpecker specimens in that museum’s collection. In particular I want to thank Steve Mlodinow, whose contagious interest in biogeography has furthered my enjoyment of birding and exploring new places.

Literature Cited:

Koenig, Walter D., Peter B. Stacey, Mark T. Stanback and Ronald L. Mumme. 1995. Acorn Woodpecker (Melanerpes formicivorus), The Birds of North America Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology; Retrieved from the Birds of North America Online: http://bna.birds.cornell.edu/bna/species/194

1

Nice summary. But Hairy Woodpecker is also found on both continents, no?

2

I consulted the South American Checklist Committe’s website (www.museum.lsu.edu/~Remsen/SACCBaseline.html) and they list only one member of the Picoides genus for South America, and that is the Smoky-brown Woodpecker (Picoides fumigatus). I love the species Latin species name — fumigatus. I was actually surprised to find that any woodpecker is native to both continents.

Speaking of Latin names, your “madaboutlutescens” handle suggests an interest in Orange-crowned Warblers, which is one of the subspecies groups I am most interested in. Here in Oregon we presume that three of the four Orange-crowned subspecies either breed in, or migrate through the state. The nominate V. c. celata is poorly documented though likely annual here as a spring and fall migrant.

3
I believe it was the British philosopher Alan Watts who said,“There are few greater kicks in life than self-righteous indignation”, While the person emoting may well give themselves a thrill or two, the main result is wasted social energy.

The battle for bio-diversity must take place on two fronts- the hearts and minds
of the emerging middle-class in the developing world; and the day to day economic reality of the billions who make up the world’s underclass.
I have been told that the primary cause of forest loss in topical areas
is not from industrial activity, but instead subsistence agriculture.
I was a guest of the Royal Projects in Thailand some years ago. The King
was sponsoring opium crop replacements. This was to protect the forests
of Thailand’s hill country as much as to reduce the world’s heroin supply.
At one experimental farm I was struck by the lack of deer fences. These
would be essential at any such operation in Oregon’s Coast Range. When
I asked the civil servant who was showing me around about this he gave
that resigned laugh I have heard too often from progressively oriented
administrators in the developing world—“There aren’t any deer in these
woods. There isn’t any wildlife of any kind. These hill people hunt every
day of the year and every hour of the day.” On our way out of the forest
I observed a large group of indigenous people gathering firewood or
the like. One of them was equipped with the most primitive firearm I
have yet to see in everyday use, probably of nineteenth century vintage.
Nonetheless, its relentless application was having very real impacts on
the local eco-system.
The current worldwide economic crisis will inevitably be felt worst
among those who had the least to start with. Economic developement
and improved standards of living doesn’t come easily, and certainly not
overnight, but that is the only solution to sustainable bio-diversity. Affluent
nations tend to do a good job of preserving their wildlife despite high
human population densities. The Netherlands for instance.

4

Nice summary and comparison, Dave. One of the angustifrons Acorn Woodpeckers I saw at San Antonia de la Sierra last year had very noticeable red on the upper breast, much like Bill’s photo from Costa Rica.

5
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